I think here the "hate" that rose against the forest is clearly meant to predate humans in the Land by quite a bit. We can also deduce that the "hate" refers to the Ravers, since the Colossus of the Fall is elsewhere stated to have been made specifically against them. It is well possible that the Ravers didn't get names or their present identities until they had met and interacted with humans, but I think they existed in some form well before that."In the nigh-unremembered past of the place which you deem the Land, the life was not the life of men and women, but of trees. One wide forest of sentience and passion filled all the region - one mind and heart alive in every leaf and bough of every tree among the many myriad throngs and the glory of the woods. And that life the Elohim loved.
"But a hate rose against the forest, seeking its destruction. And this was dire, for a tree may love and feel pain and cry out, but has few means of defense. The knowledge was lacking. Therefore we met, and from among us Appointed one to give her life to that forest. This she did by merging among the trees until they gained the knowledge they required.
"Their knowledge they employed to bind her in stone, exercising her name and being to form an interdict against that hate. Thus was she lost to herself and to her people, but the interdict remained while the will of the forest remained to hold it."
"The Colossus," Covenant breathed. "The Colossus of the Fall."
"Yes," Findail said.
"And when people started coming to the Land, started cutting down the trees as if they were just so much timber and difficulty, the forest used what it'd learned to create the Forestals in self-defense. Only it took too long, and there were too many people, and the Forestals weren't enough, they couldn't be everywhere at once, couldn't stop the many blind or cruel or simply unscrupulous axes and fires. They were lucky to keep the mind of the forest awake as long as they did."
"Yes," Findail said again.
I'm worried that we may have a continuity error here... Between Findail and Anele I'm however inclined to believe Findail has it right. Neither character has any reason to lie, but Findail was actually alive at the time and peripherally involved, while Anele is relying on second-hand information. Rock experiences time even slower than trees, so that it might miss fleeting things, and it is unknown if fracturing has a detrimental effect on the integrity of stony memories.